R. J. Scholes, The Influence of Soil Fertility on the Ecology of Southern African Dry Savannas, Journal of Biogeography, Vol. 17, No. 4/5, Savanna Ecology and Management: Australian Perspectives and Intercontinental Comparisons (Jul. - Sep., 1990), pp. 415-419
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Robert J. Scholes (1990) studied this question.