P. Dennis, M. R. Young, C. L. Howard, I. J. Gordon, The Response of Epigeal Beetles (Col.: Carabidae, Staphylinidae) to Varied Grazing Regimes on Upland Nardus stricta Grasslands, Journal of Applied Ecology, Vol. 34, No. 2 (Apr., 1997), pp. 433-443
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Dennis et al. (1997) studied this question.
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