M. C. S. Kingsley, J. A. Nagy, R. H. Russell, Patterns of Weight Gain and Loss for Grizzly Bears in Northern Canada, Bears: Their Biology and Management, Vol. 5, A Selection of Papers from the Fifth International Conference on Bear Research and Management, Madison, Wisconsin, USA, February 1980 (1983), pp. 174-178
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Kingsley et al. (1983) studied this question.
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