G. L. Vourlitis, W. C. Oechel, S. J. Hastings, M. A. Jenkins, A System for Measuring in situ CO2 and CH4 Flux in Unmanaged Ecosystems: An Arctic Example, Functional Ecology, Vol. 7, No. 3 (Jun., 1993), pp. 369-379
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Vourlitis et al. (1993) studied this question.