W. E. Castle, F. W. Carpenter, A. H. Clark, S. O. Mast, W. M. Barrows, The Effects of Inbreeding, Cross-Breeding, and Selection upon the Fertility and Variability of Drosophila, Proceedings of the American Academy of Arts and Sciences, Vol. 41, No. 33 (May, 1906), pp. 731-786
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Castle et al. (1906) studied this question.