nanometre- thin films of adhesive fluid. This fluid helps to increase the pad's contact area on rough surfaces and its special rheology combines capillary adhesion with resistance against sliding. We summarize recent advances in the understanding of fluid-based insect adhesion and highlight the large number of unresolved questions, regarding the mechanism of fluid secretion, the thickness of the mediating fluid layer, and the influence of surface properties on insect adhesive forces.
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Dirks et al. (2011) studied this question.
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