Simon P. Penson, Robert C. Schuurink, Angelika Fath, Frank Gubler, John V. Jacobsen, Russell L. Jones, cGMP Is Required for Gibberellic Acid-Induced Gene Expression in Barley Aleurone, The Plant Cell, Vol. 8, No. 12 (Dec., 1996), pp. 2325-2333
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Penson et al. (1996) studied this question.