J. D. Goss-Custard, R. W. G. Caldow, R. T. Clarke, S. E. A. le V. dit Durell, W. J. Sutherland, Deriving Population Parameters from Individual Variations in Foraging Behaviour. I. Empirical Game Theory Distribution Model of Oystercatchers Haematopus ostralegus Feeding on Mussels Mytilus edulis, Journal of Animal Ecology, Vol. 64, No. 2 (Mar., 1995), pp. 265-276
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Goss‐Custard et al. (1995) studied this question.