The rate parameters R1, R2, I/LI and I/t0.5, which characterize the growth in area of successive main-stem leaves, probably all have the same temperature response. Temperature therefore only operates on the time scale. Water stress reduces both the relative growth rate and the advance of developmental age, the latter however to a lesser extent than the former. The effect of root restriction is explained as resulting from mineral shortage.
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H.J.W. Mutsaers (1983) studied this question.
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