Three new species segregated from Phebalium squamulosum subsp. squamulosum (Rutaceae) based on morphological and phytochemical data

The genus PhebaliumVen t. is widespread through southern and eastern Australia with 28 species recognised in the 'Flora of Australia' (Wilson 2013) and 30 recognised by APC (2016). Several species, in particular Phebalium squamulosum Vent., exhibit considerable morphological variation. In the last major revision of the genus, Telford and Bruhl (2014) subsequently raised P. squamulosum subsp. verrucosum Paul G.Wilson to the rank of species. The type specimen of P. squamulosum was taken from a plant cultivated in Empress Josephine's garden at Malmaison, France, from a collection from the Sydney area,"originaire de la Nouvelle Galles, et croissant sur les montagnes" (Ventenat 1805). A lectotype was selected by Callmander et al. (2017), which includes an image of an isolectotype. In his treatment of P. squamulosum subsp. squamulosum, Wilson (1970) included lowland plants of the Sydney Basin as well as plants from south eastern Queensland and high altitudes of the New England Tableland, Abstract Phebalium graniticola I.Telford & JJ.Bruhl, P. stellatum I.Telford & JJ.Bruhl and P. sylvaticum I.Telford & JJ.Bruhl are segregated as new species from Phebalium squamulosum Vent, subsp.