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The aim of the investigation was to find a method of growing birch seedlings with a constant optimum nutrient status. The studies in this paper have indicated important factors in such a growth method which must be comprised in a definition of nutrient requirements. The optimum nutrient proportions in the seedlings and the optimum total concentration in the solution must be known. Special attention must be paid to the ratio between NH 4 + and NO 3 − in the solution and in the uptake. The amounts of nutrients taken up at optimum nutrition can be followed by measurement of pH and conductivity. Thus, the nutrients consumed may be replaced by titrations which may be automated by standard equipment. The nutrient proportions to be added are determined by the optimum internal proportions. It is not necessary to change the nutrient solution even if the volume is relatively small in relation to plant mass. There was no evidence of effects of accumulation of root exudates or infections. By using the system described, it is possible to attain considerably greater fresh and dry matter production in birch seedlings than when an optimum conventional solution is used.
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Torsten Ingestad (1970) studied this question.
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