Barton D. Clinton, Lindsay R. Boring, Wayne T. Swank, Regeneration Patterns in Canopy Gaps of Mixed-Oak Forests of the Southern Appalachians: Influences of Topographic Position and Evergreen Understory, The American Midland Naturalist, Vol. 132, No. 2 (Oct., 1994), pp. 308-319
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Clinton et al. (1994) studied this question.
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