Douglas E. Brash, Ronald W. Hart, Biopsy Measurement of DNA Damage and Repair in Vivo: Single-Strand Breaks, Alkali-Labile Bonds, and Endonuclease-Sensitive Sites, Radiation Research, Vol. 91, No. 1 (Jul., 1982), pp. 169-180
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Brash et al. (1982) studied this question.