A. C. Blackmore, M. T. Mentis, R. J. Scholes, The Origin and Extent of Nutrient-Enriched Patches within a Nutrient-Poor Savanna in South Africa, Journal of Biogeography, Vol. 17, No. 4/5, Savanna Ecology and Management: Australian Perspectives and Intercontinental Comparisons (Jul. - Sep., 1990), pp. 463-470
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Blackmore et al. (1990) studied this question.
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