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Gabriel Neve, Bernard Barascud, Rosemary Hughes, Josiane Aubert, Henri Descimon, Philippe Lebrun, Michel Baguette, Dispersal, Colonization Power and Metapopulation Structure in the Vulnerable Butterfly Proclossiana eunomia (Lepidoptera: Nymphalidae), Journal of Applied Ecology, Vol. 33, No. 1 (Feb., 1996), pp. 14-22
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Nève et al. (1996) studied this question.
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