Isobel Radforth, The Food of the Grayling (Thymallus thymallus), Flounder (Platichthys flesus), Roach (Rutilus rutilus) and Gudgeon (Gobio fluviatilis), with Special Reference to the Tweed Watershed, Journal of Animal Ecology, Vol. 9, No. 2 (Nov., 1940), pp. 302-318
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Isobel Radforth (1940) studied this question.
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