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Neil Snow, The Phylogenetic Utility of Lemmatal Micromorphology in Leptochloa S. L. an Related Genera in Subtribe Eleusininae (Poaceae, Chloridoideae, Eragrostideae), Annals of the Missouri Botanical Garden, Vol. 83, No. 4 (1996), pp. 504-529
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Neil Snow (1996) studied this question.
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