Anja G. J. Kuipers, Evert Jacobsen, Richard G. F. Visser, Formation and Deposition of Amylose in the Potato Tuber Starch Granule Are Affected by the Reduction of Granule-Bound Starch Synthase Gene Expression, The Plant Cell, Vol. 6, No. 1 (Jan., 1994), pp. 43-52
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Kuipers et al. (1994) studied this question.