N. Leader-Williams, S. D. Albon, P. S. M. Berry, Illegal Exploitation of Black Rhinoceros and Elephant Populations: Patterns of Decline, Law Enforcement and Patrol Effort in Luangwa Valley, Zambia, Journal of Applied Ecology, Vol. 27, No. 3 (Dec., 1990), pp. 1055-1087
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Leader‐Williams et al. (1990) studied this question.
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