Spermatophore formation and transfer in slugs are complex and still obscure. In the slug Arion lusitanicus Mab. the spermatophore is formed 30-35 minutes after atrium eversion; as long as the copulating slugs are connected, each spermatophore is partly in the donor's epiphallus and partly in the recipient's spermatheca duct; spermatozoa are released in the spermatheca duct 12-13 hrs after the spermatophore transfer; the spermatophore tunicle in the spermatheca dissolves within 48-72 hrs.
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Sionek et al. (2001) studied this question.
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