K. P. Currens, C. B. Schreck, H. W. Li, Allozyme and Morphological Divergence of Rainbow Trout (Oncorhynchus mykiss) above and below Waterfalls in the Deschutes River, Oregon, Copeia, Vol. 1990, No. 3 (Sep. 19, 1990), pp. 730-746
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Currens et al. (1990) studied this question.
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