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N. J. Mills, A. P. Gutierrez, Prospective Modelling in Biological Control: An Analysis of the Dynamics of Heteronomous Hyperparasitism in a Cotton-Whitefly-Parasitoid System, Journal of Applied Ecology, Vol. 33, No. 6 (Dec., 1996), pp. 1379-1394
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Mills et al. (1996) studied this question.
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