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Richard Borowsky, Klaus D. Kallman, Patterns of Mating in Natural Populations of Xiphophorus (Pisces: Poeciliidae). I: X. maculatus from Belize and Mexico, Evolution, Vol. 30, No. 4 (Dec., 1976), pp. 693-706
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Borowsky et al. (1976) studied this question.
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