G. H. Whitesides, J. F. Oates, S. M. Green, R. P. Kluberdanz, Estimating Primate Densities from Transects in a West African Rain Forest: A Comparison of Techniques, Journal of Animal Ecology, Vol. 57, No. 2 (Jun., 1988), pp. 345-367
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Whitesides et al. (1988) studied this question.
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