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Robert W. Hoshaw, Charles V. Wells, Systematics of the Zygnemataceae (Chlorophyceae). IV. Nuclear Cytology of Sirogonium melanosporum, a Species with Large Chromosomes, Transactions of the American Microscopical Society, Vol. 101, No. 3 (Jul., 1982), pp. 276-286
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Hoshaw et al. (1982) studied this question.
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