Mark Leighton, Donna R. Leighton, The Relationship of Size of Feeding Aggregate to Size of Food Patch: Howler Monkeys (Alouatta palliata) Feeding in Trichilia cipo Fruit Trees on Barro Colorado Island, Biotropica, Vol. 14, No. 2 (Jun., 1982), pp. 81-90
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Leighton et al. (1982) studied this question.