Composition and size of groups arc (bought to have evolved as adapta-tions that increase efficiency in exploiting and defending food resources and that reduce the risk of predations (BERTRAM, 1978; DUNBAR, 1988; TERBORGH, 1983; TERBORGH & JANSON, 1986). Disagreements persist
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Christophe Boesch (1991) studied this question.
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