F. S. Chapin III, J. D. McKendrick, D. A. Johnson, Seasonal Changes in Carbon Fractions in Alaskan Tundra Plants of Differing Growth Form: Implications for Herbivory, Journal of Ecology, Vol. 74, No. 3 (Sep., 1986), pp. 707-731
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Chapin et al. (1986) studied this question.
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