B. Jedrzejewska, H. Okarma, W. Jedrzejewski, L. Milkowski, Effects of Exploitation and Protection on Forest Structure, Ungulate Density and Wolf Predation in Bialowieza Primeval Forest, Poland, Journal of Applied Ecology, Vol. 31, No. 4 (Nov., 1994), pp. 664-676
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Jędrzejewska et al. (1994) studied this question.
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