David A. Erasmus, The Host-Parasite Interface of Cyathocotyle bushiensis Khan, 1962 (Trematoda: Strigeoidea) II. Electron Microscope Studies of the Tegument, The Journal of Parasitology, Vol. 53, No. 4 (Aug., 1967), pp. 703-714
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David A. Erasmus (1967) studied this question.