Ken Matsuoka, Tatsuji Higuchi, Masayoshi Maeshima, Kenzo Nakamura, A Vacuolar-Type H⁺ ATPase in a Nonvacuolar Organelle Is Required for the Sorting of Soluble Vacuolar Protein Precursors in Tobacco Cells, The Plant Cell, Vol. 9, No. 4 (Apr., 1997), pp. 533-546
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Matsuoka et al. (1997) studied this question.