I. S. Darevsky, Natural Parthenogenesis in a Polymorphic Group of Caucasian Rock Lizards Related to Lacerta saxicola Eversmann, Journal of the Ohio Herpetological Society, Vol. 5, No. 4 (Dec. 30, 1966), pp. 115-152
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Ilya S. Darevsky (1966) studied this question.