C. Thomas Philbrick, Gregory J. Anderson, Implications of Pollen/Ovule Ratios and Pollen Size for the Reproductive Biology of Potamogeton and Autogamy in Aquatic Angiosperms, Systematic Botany, Vol. 12, No. 1 (Jan. - Mar., 1987), pp. 98-105
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Philbrick et al. (1987) studied this question.
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