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Marguerite J. Varagona, Robert J. Schmidt, Natasha V. Raikhel, Nuclear Localization Signal(s) Required for Nuclear Targeting of the Maize Regulatory Protein Opaque-2, The Plant Cell, Vol. 4, No. 10 (Oct., 1992), pp. 1213-1227
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Varagona et al. (1992) studied this question.
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