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Gwyneth C. Ingram, Justin Goodrich, Mark D. Wilkinson, , George W. Haughn, Enrico S. Coen, Parallels between Unusual Floral Organs and FIMBRIATA, Genes Controlling Flower Development in Arabidopsis and Antirrhinum, The Plant Cell, Vol. 7, No. 9 (Sep., 1995), pp. 1501-1510
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Ingram et al. (1995) studied this question.
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