M. O. Hill, R. G. H. Bunce, M. W. Shaw, Indicator Species Analysis, A Divisive Polythetic Method of Classification, and its Application to a Survey of Native Pinewoods in Scotland, Journal of Ecology, Vol. 63, No. 2 (Jul., 1975), pp. 597-613
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Hill et al. (1975) studied this question.
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