J. P. Lumaret, E. Galante, C. Lumbreras, J. Mena, M. Bertrand, J. L. Bernal, J. F. Cooper, N. Kadiri, D. Crowe, Field Effects of Ivermectin Residues on Dung Beetles, Journal of Applied Ecology, Vol. 30, No. 3 (1993), pp. 428-436
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Lumaret et al. (1993) studied this question.