In the tortoise, Gopherus polyphemus, single unit spikes in the olfactory epithelium in response to amyl acetate were positive relative to the slow potential. The number of spikes in a response train was 4 to 15, the duration 3 to 4 msec, the height 0.5 to 2 mv. The height of successive spikes in a train decreased. The decrement in height, the number, and the frequency of spikes changed with the strength of the odor.
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Shibuya et al. (1963) studied this question.
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