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Two distinct nucellar cell types occur in barley: rectangular epidermal cells which persist through early embryogenesis, and irregular, highly vacuolate interior cells which seem to undergo lysis as the embryo and endosperm develop Both types have lobed nuclei. Breakdown of the nucellus results in formation of a layer of nucellar debris termed the "nucellar lysate." This layer surrounds the endosperm and may be absorbed by the endosperm during its development or may contribute to the development of the aleurone wall.
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Knut Norstog (1974) studied this question.
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