Like a complicated puzzle containing pieces that do not belong, the determination of the structure of the active site of photosystem II presents several intricacies. The picture on the right shows a new model for the oxo- and carboxylato-bridged Mn4 cluster, which also considers the coordinated cofactors Ca2+ and Cl−. The concept is based on spectroscopic data (ESR, EXAFS) from years of research.
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Karl Wieghardt (1994) studied this question.
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