John C. Avise, Michael H. Smith, Robert K. Selander, Timothy E. Lawlor, Paul R. Ramsey, Biochemical Polymorphism and Systematics in the Genus Peromyscus. V. Insular and Mainland Species of the Subgenus Haplomylomys, Systematic Zoology, Vol. 23, No. 2 (Jun., 1974), pp. 226-238
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Avise et al. (1974) studied this question.