A recent study of the ecological and reproductive relationships between Aquilegia formosa and A. pubescens (Ranunculaceae) in the Sierra Nevada of California My 1952 paper dealt with several interrelated topics: ecological isolation, floral isolation, and introgression between the two species or semispecies A. formosa and A. pube.scens; and, in the light of this, the basic species groups and mode of adaptive radiation in the genus Aquilegia. The objective of The differences between the recent and the earlier treatments are numerous, and range from significant to trivial issues.
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Verne Grant (1976) studied this question.
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