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In order to study the ecology of intertidal plants and animals it is desirable to choose a coast-line which is as simple as possible. When numerous factors vary simultaneously it is impossible to decide which of them is responsible for any change which may occur in the flora and fauna. The complexity of a shore which is ravined or boulder-strewn makes ecological work difficult. Unfortunately, certain major influences, such as wave action, involve a whole complex of subsidiary effects, so that the perfect locality, where each environmental factor would be found to vary in turn independently of all the other factors, cannot exist.
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J. A. Kitching (1935) studied this question.
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