I. J. Odrzykoski, L. D. Gottlieb, Duplications of Genes Coding 6-Phosphogluconate Dehydrogenase in Clarkia (Onagraceae) and Their Phylogenetic Implications, Systematic Botany, Vol. 9, No. 4 (Oct. - Dec., 1984), pp. 479-489
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Odrzykoski et al. (1984) studied this question.
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