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Peter J. Grubb, William G. Lee, Johannes Kollmann, J. Bastow Wilson, Interaction of Irradiance and Soil Nutrient Supply on Growth of Seedlings of Ten European Tall-Shrub Species and Fagus Sylvatica, Journal of Ecology, Vol. 84, No. 6 (Dec., 1996), pp. 827-840
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Grubb et al. (1996) studied this question.
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