Charles Atwood Kofoid, Olive Swezy, Mitosis and Multiple Fission in Trichomonad Flagellates, Proceedings of the American Academy of Arts and Sciences, Vol. 51, No. 6 (Nov., 1915), pp. 289-364, 366, 368, 370, 372, 374, 376, 378
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Kofoid et al. (1915) studied this question.