Guarino R. Colli, Alexandre F. B. de Araújo, Ronis da Silveira, Fernando Roma, Niche Partitioning and Morphology of Two Syntopic Tropidurus (Sauria: Tropiduridae) in Mato Grosso, Brazil, Journal of Herpetology, Vol. 26, No. 1 (Mar., 1992), pp. 66-69
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Colli et al. (1992) studied this question.