C. G. T. Morison, A. C. Hoyle, J. F. Hope-Simpson, Tropical Soil-Vegetation Catenas and Mosaics: A Study in the South-Western Part of the Anglo-Egyptian Sudan, Journal of Ecology, Vol. 36, No. 1 (Jul., 1948), pp. 1-84
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Morison et al. (1948) studied this question.