Authors
M. L. Fernald, Persistence of Plants in Unglaciated Areas of Boreal America, Memoirs of the American Academy of Arts and Sciences, New Series, Vol. 15, No. 3, Persistence of Plants in Unglaciated Areas of Boreal America (Jun. 25, 1925), pp. 239, 241-342
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Merritt Lyndon Fernald (1925) studied this question.