Thomas A. Ranker, Christopher H. Haufler, Pamela S. Soltis, Douglas E. Soltis, Genetic Evidence for Allopolyploidy in the Neotropical Fern Hemionitis pinnatifida (Adiantaceae) and the Reconstruction of an Ancestral Genome, Systematic Botany, Vol. 14, No. 4 (Oct. - Dec., 1989), pp. 439-447
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Ranker et al. (1989) studied this question.
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